Antumbra and Aspen's youngest daughter from their first litter and Moonchaser's great-great-great-granddaughter, Willow has been chosen as the next pack leader after Dewdrop, Snowdrop, Borealis and Antumbra all turned the role down in favor of other roles. Willow was the only wolf who truly wanted the responsibility of leading the pack. Despite her adventurous spirit, she isn't reckless. She's been craving to be a useful member of the Wolves of The Willows pack for a long time now, and what better way to assist her packmates by being a responsible and welcoming role model? She is more temperamental than Moonchaser, though, sometimes being grouchy when other animals annoy her. She tries not to get annoyed at her packmates, though, because Moonchaser advises her successor that "Annoyance and hate can tear relationships apart", and the last thing that Willow wants is for the pack she grew up in to fall apart. She's become smitten with the sheepish and sensitive Ginkgo, who will take Glacier's place as second alpha and lead pupsitter due to his position as Willow's mate.