Tioga and Nittany were one of the first wolves to join Dioute at Wapiti Run. They were both older wolves who lived in the area and wanted more than just the lone wolf life. They settled in easily and provided a nurturing environment to those who joined.
However, Tioga was always more of a feisty one, quick to say exactly what she thought and exactly what was on her mind. She is close with her hunting patrol, Maren, Nittany, Sutha (deceased) and the pupsitter, Doguinet.