Leto doesn't know who his biological parents are, but he dose know his adopted parents well....
Leto's father, Felt, was mighty hunter of the pack and had an close relationship to the Alpha, Armonk, of the pack and would eventually become mates with the Beta, Chiro.
Chiro and Felt would have many pups with only 5 being noticeable, Tiarnán, Elsdon, Jamshid, Elisha, and Loreen. Leto and Har was also a noticeable pups of theirs but they were not blood related to Chiro and Felt...
Tiarnán was almost exactly like his father his only difference being that he had his moms coat, he was also a great hunter.
Elsdon was all bronze no brains and made for a good hunter if paired with his older and twin brothers.
Jamshid is Elsdon twin brother and the complete opposite of him, with only there size and finesse being the same.
Elisha was the smallest and quickest pup of the herd, and as such had apprentice under the greatest scout of the pack and slowly toke his place.
Loreen was the odd one out of the 5 pups, for one she was a girl and second she was able to topple all of her brothers including Elsdon and Tiarnán. Although she would have made for a great hunter or scout she decided to follow in her mother's footstep and become the next herbalist. Cause of this her and Leto became close.
Leto was found by Chiro next to an river, barely two weeks old almost frozen and basically skin and bones, she would later say that see saw something in him and cause of that she toke him and nursed him alone with the latest litter of pups she had.
Heiko Yelisey
Herbalist 
Herbalist