Spruce and his two sisters Wisteria and Sycamore were born among humans, they were a part of a rehabilitation program for wolves. though Sycamore hated humans with a burning passion Spruce felt at least something for the creatures who had cared for him and his littermates since they were born. but eventually the humans started to come less and less leaving the wolves to their own devices for months supplying food ever so often. the three siblings were forced to defend themselves against the other wolves which led to Spruce being half blind. eventually new humans arrived, these ones put a weird collar on Spruce and Wisteria and before they knew it they woke up in the forest alone with no humans and were forced to survive.